Biswas, S. and Chatterjee, S.K. 1984. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India. 82, 1-4 (Dec. 1984), 147–177. DOI:https://doi.org/10.26515/rzsi/v82/i1-4/1984/161291.